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Amber Tamblyn quickly earned widespread critical acclaim and a Golden Globe Award nomination for Best Dramatic Actress in a Drama Series for her portrayal of Joan during last year's premiere season of JOAN OF ARCADIA. She had previously been best known for playing Emily Quartermaine in the daytime drama "General Hospital." That role, which she originated in 1995, was set to last for only a few months, but it turned into a seven-year stint as she won viewers' hearts, critics' acclaim and two consecutive Hollywood Reporter Young Star Awards as Best Young Actress in a Daytime Series.

Tamblyn's most recent film credit is the thriller "The Ring." Her future film projects include a lead role in "Sisterhood of the Traveling Pants," an adaptation of the Ann Brashares novel about four friends who keep in touch by circulating a magical pair of secondhand jeans.

Her additional primetime television credits include guest-starring roles in "Buffy the Vampire Slayer," "The New Twilight Zone" and "Boston Public," as well as CSI: MIAMI and WITHOUT A TRACE on CBS. She also appeared in the short film "Ten Minutes Older: The Trumpet."

Her father is veteran actor Russ Tamblyn ("West Side Story," "Seven Brides for Seven Brothers," "Tom Thumb" and "Twin Peaks"). She was born and raised in Los Angeles, where she lives. Her birth date is May 14.

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Joe Mantegna

as Will Girardi

Mary Steenburgen

as Helen Girardi

Amber Tamblyn

as Joan Girardi

Jason Ritter

as Kevin Girardi

Michael Welch

as Luke Girardi

Christopher Marquette

as Adam Rove

Becky Wahlstrom

as Grace Polk